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The second film in a duology of Uchuu Senkan Yamato 2205: Aratanaru Tabidachi.

In the year 2000, Alice Hayashi wins a trip to the moon in a sweepstake. However, when her shuttle launches, something goes terribly wrong, and the next thing she knows, she has crashed somewhere in a frozen wasteland. Before she can get her bearings, strange soldiers begin to attack her, forcing her to flee. Eventually, Alice meets a young man called Yuan, who tells her that she is in Lapland in the year 2030. The soldiers pursuing her belong to Nero, an enigmatic leader who controls the world through his supercomputer. He is also responsible for reducing the Earth's population by over seven billion. Convinced that Nero has something to do with her time jump, Alice, Yuan, and the android maid Maria journey to Kingdom, the shadowy fortress that Nero rules from.

In the year 2000, Alice Hayashi wins a trip to the moon in a sweepstake. However, when her shuttle launches, something goes terribly wrong, and the next thing she knows, she has crashed somewhere in a frozen wasteland. Before she can get her bearings, strange soldiers begin to attack her, forcing her to flee. Eventually, Alice meets a young man called Yuan, who tells her that she is in Lapland in the year 2030. The soldiers pursuing her belong to Nero, an enigmatic leader who controls the world through his supercomputer. He is also responsible for reducing the Earth's population by over seven billion. Convinced that Nero has something to do with her time jump, Alice, Yuan, and the android maid Maria journey to Kingdom, the shadowy fortress that Nero rules from.

After the Earth was destroyed by mysterious alien lifeforms known as the Gauna, surviving remnants of the human race escaped to space in the enormous generation ship Sidonia. Having drifted through space for millennia, the Sidonia found itself once more under attack from Gauna for the first time in a century. Once again facing the threat of extinction, a temporary victory against the Gauna was eked out thanks to the human-Gauna hybrid Tsumugi Shiraui and ace Guardian mech pilot Nagate Tanikaze. 10 years later... The people of Sidonia enjoy a brief respite. Peaceful days pass, during which Tsumugi begins to realize her feelings for Nagate, who is now celebrated as a hero of Sidonia. However, as Captain Kobayashi has always known, as long as the Gauna remain, peace cannot last. The decision is made: a final battle, upon which rests the fate of humanity's last survivors. As the end approaches, will the crew be able to protect those they love?